蹄盖蕨属
水龙骨目、蹄盖蕨科下级分类
蹄盖蕨属,学名:Athyrium Roth,为水龙骨目、蹄盖蕨科下级分类。
形态特征
植物形态
陆生,中型草本植物。根状茎短,多为直立,少有横走或斜升,罕细长而横走;叶簇生,罕近生或远生。
叶子形态
叶柄长,基部往往加粗,背面隆起腹面凹陷,两侧边缘有瘤状气囊体各一行,向下变为尖削,少为圆柱形,有时无气囊体,向下不尖削;横切面有2条维管束,呈八字形排列,向叶轴连合呈U字形;叶柄基部被鳞片,鳞片红褐色、深褐色或黑褐色,卵状披针形、长钻形或线状披针形,全缘,由厚壁的狭长细胞构成,膜质,基部着生;叶柄上面有1条纵沟,向上直通叶轴,沟内往往被短腺毛;叶片卵形、长圆形或阔披针形,一回至三回羽状,各回羽片基部合生或分离,通常以锐角沿各回羽轴下面圆形,上面有1条深纵沟,两侧边缘呈刀口状的隆起,但与其上一回羽轴(或主脉)汇合处有1断裂缺口,彼此互通,其下无或有1个肉质的刺状凸起。叶脉分离,叉状或羽状,小脉伸达锯齿顶端。叶干后常为草质,少为纸质或革质,两面通常光滑,罕被鳞片或毛,唯叶轴、羽轴和小羽轴(或主脉)上往往被有短腺毛。
孢子形态
孢子囊群圆形、圆肾形、马蹄形、弯勾形、长圆形或短线形,背生、侧生或横跨小脉上;囊群盖圆肾形、马蹄形、弯勾形、新月形、长圆形或短线形,褐色,膜质,边缘啮蚀状或有睫毛,少为全缘,通常宿存,罕无囊群盖或囊群盖不发育。孢子两面型,极面观为椭圆形,赤道面观为豆形,周壁表面具褶皱或无褶皱。
染色体
染色体基数x=40
植株特点
杂交是造成本属分类困难的原因之一,在日本已报道有60多个杂种,中国究竟有多少杂种还有待进一步研究。
属的模式种:Athyrium filix-femina (L.) Roth——Polypodium filix-femina L.
下级组
蹄盖蕨组Sect. Athyrium S. K. Wu et R. C. Ching
疏叶蹄盖蕨组Sect. Dissitifolia (X. C. Zhang) Z. R. Wang
海南蹄盖蕨组Sect. Hainanensia Z. R. Wang
川滇蹄盖蕨组Sect. Mackinnoniana (Ching et Y. T. Hsieh) Z. R. Wang
小蹄盖蕨组Sect. Minima Z. R. Wang
多羽蹄盖蕨组Sect. Multipinna Z. R. Wang
日本蹄盖蕨组Sect. Niponica Ching et Y. T. Hsieh
刺蹄盖蕨组Sect. Otophora Z. R. Wang
裸囊蹄盖蕨组Sect. Pachyphylla Z. R. Wang
篦齿蹄盖蕨组Sect. Pectinata (Ching et Hsieh) Z. R. Wang
耳状蹄盖蕨组Sect. Polystichoides Ching et Y. T. Hsieh
狭基蹄盖蕨组Sect. Stenopoda Z. R. Wang
毛叶蹄盖蕨组Sect. Suprapubescentia Z. R. Wang
黑秆蹄盖蕨组Sect. Wallichiana (X. C. Zhang) Z. R. Wan
下级物种
原生物种
自然杂交物种
Athyrium × akiense Sa.Kurata
Athyrium × amagi-pedis Sa.Kurata
Athyrium × anceps Sa.Kurata
Athyrium × austrojaponense Sa.Kurata
Athyrium × awatae Seriz.
Athyrium × bicolor Seriz.
Athyrium × boreo-occidentali-indobharaticola-birianum Fraser-Jenk.
Athyrium × calophyllum Sa.Kurata ex Seriz.
Athyrium × cornopteroides Sa.Kurata
Athyrium × decurrentiserratum K.Hori
Athyrium × dixitii Fraser-Jenk.
Athyrium × drepanopteroides Fraser-Jenk.
Athyrium × flavosorum Seriz.
Athyrium × fuscopaleaceum Sa.Kurata
Athyrium × glabrescens Seriz.
Athyrium × gurungae Fraser-Jenk.
Athyrium × hisatsuanum Sa.Kurata
Athyrium × ikutae Sa.Kurata
Athyrium × inabaense Seriz.
Athyrium × inouei Sa.Kurata
Athyrium × kathmanduense Fraser-Jenk.
Athyrium × kawabatae Sa.Kurata
Athyrium × kawabataeoides Sa.Kurata
Athyrium × kidoanum Sa.Kurata
Athyrium × langtangense Fraser-Jenk.
Athyrium × lantangense Fraser-Jenk.
Athyrium × lobulosoimpolitum Fraser-Jenk.
Athyrium × manickamii Fraser-Jenk.
Athyrium × mantoniae Fraser-Jenk.
Athyrium × masachikanum (Sa.Kurata) Seriz.
Athyrium × masayukianum Sa.Kurata
Athyrium × megayakusimense Sa.Kurata
Athyrium × meghalaicum Fraser-Jenk.
Athyrium × mentiens Sa.Kurata
Athyrium × minakuchii Sa.Kurata
Athyrium × multifidum Rosenst.
Athyrium × neoelegans Sa.Kurata
Athyrium × nepalense Fraser-Jenk.
Athyrium × paludicola Sa.Kurata ex Seriz.
Athyrium × petiolulatum Sa.Kurata
Athyrium × pichisermollianum Fraser-Jenk.
Athyrium × pokharense Fraser-Jenk.
Athyrium × pseudobipinnatum K.Hori
Athyrium × pseudoiseanum Sa.Kurata
Athyrium × pseudopinetorum Seriz.
Athyrium × pseudospinescens Seriz.
Athyrium × pseudowardii Seriz.
Athyrium × purpurascens (Tagawa) Sa.Kurata
Athyrium × purpureipes Sa.Kurata
Athyrium × pygmaei-silvae Sa.Kurata
Athyrium × quaesitum Sa.Kurata
Athyrium × reichsteinii Schneller & Rasbach
Athyrium × saitoanum (Sugim.) Seriz.
Athyrium × sanjappae Fraser-Jenk.
Athyrium × satsumense Seriz.
Athyrium × sefuricola Sa.Kurata
Athyrium × shiibaense (Seriz.) Nakaike
Athyrium × shikoku-montanum Seriz.
Athyrium × simlense Fraser-Jenk.
Athyrium × sledgei Fraser-Jenk.
Athyrium × subchristensenianum K.Hori
Athyrium × subcrassipes Seriz.
Athyrium × tokashikii Sa.Kurata
Athyrium × tsurutanum Sa.Kurata
Athyrium × undulatipinnulum Seriz.
Athyrium × watanabei Seriz.
Athyrium × yakuinsulare Sa.Kurata
Athyrium × yakumonticola Sa.Kurata
异型异名
Anisocampium C.Presl in Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 58 (1851)
× Apseudathyrium Xia Wan & Li Bing Zhang in Phytotaxa 575: 104 (2022)
× Cornoathyrium Nakaike in New Fl. Jap. Pterid.: 840 (1992)
Cornopteris Nakai in Bot. Mag. (Tokyo) 44: 7 (1930)
Ephemeropteris R.C.Moran & Sundue in Taxon 68: 432 (2019)
Kuniwatsukia Pic.Serm. in Webbia 28: 455 (1973)
Microchlaena Ching in Bull. Fan Mem. Inst. Biol., Bot. 8: 322 (1938), nom. illeg.
Neoathyrium Ching & Z.R.Wang in Acta Phytotax. Sin. 20: 76 (1982)
Oligocampia Trevis. in Atti Reale Ist. Veneto Sci. Lett. Arti, ser. 2, 2: 165 (1851)
Pseudathyrium Newman in Phytologist 4: 370 (1851)
Pseudocystopteris Ching in Acta Phytotax. Sin. 9: 76 (1964)
× Schnellerathyrium Hong M.Liu, Schuettp. & H.Schneid. in J. Syst. Evol. 58: 999 (2020), not validly publ.
最新修订时间:2024-04-01 19:12
目录
概述
形态特征
参考资料